Founded in 1988

The Mara Hyena Project was co-founded by Dr. Kay Holekamp and Dr. Laura Smale of Michigan State University on May 15, 1988. What began as a three-year study to understand how female spotted hyenas came to dominate males grew into one of the longest continuously running wildlife studies in Africa.

From the outset, the project established a commitment to individual-level data collection, identifying every hyena in the study clan by spot pattern and ear notches, and recording detailed behavioral observations on each animal from birth through adulthood. This longitudinal approach is what makes the MHP dataset uniquely powerful.

Dr. Kay Holekamp in the field

May 15, 1988

Project Founded

Dr. Kay Holekamp and Dr. Laura Smale arrive in the Maasai Mara and begin the study, initially focusing on a single hyena clan near Talek. Kay and Laura focused on slowly habituating hyenas to the presence of research vehicles, a process that laid the groundwork for everything that followed.

At this time, the study focused on just one clan of roughly 50 hyenas, and Talek town was a small settlement of only a few hundred people with no school or hospital. Local Maasai community members would often seek Kay's help when they needed medical attention, calling her Mama Fisi, meaning hyena lady, and Kay would do what she could or drive them to the hospital in Narok.

Dr. Kay Holekamp in the early field years
Dr. Kay Holekamp
Dr. Laura Smale in the early field years
Dr. Laura Smale

1988 onwards

One Question Becomes Many

What started as a single question about female dominance quickly gave rise to a whole new set of questions: What happens when male offspring leave their natal clans and disperse? What shape does hyena intelligence take? Does rank at birth affect adult outcomes? Each answer opened new avenues of inquiry that continue to drive MHP research today.

1992

Kay and Laura Join MSU Faculty

Dr. Holekamp and Dr. Smale both join the faculty of Michigan State University, establishing the institutional home the MHP has maintained ever since. Dr. Smale subsequently founded an independent neuroscience lab at MSU, while Kay continued leading the hyena project. Laura remained closely connected to the project, serving on the PhD committees of many of Kay's students over the following decades.

Dr. Kay Holekamp

2008

Second Field Site Opens at Serena

The project expands to a second field camp near Serena Lodge, adding the Serena North, Serena South, and HZ clans to the study. This expansion provides a spatial comparison across the reserve and substantially increases the scope of long-term data collection.

2010

Malit Benson Pion Joins the Project

Malit Benson Pion grew up herding cattle in the Talek region and as a boy would hide from the MHP's research vehicles, mistaking them for ranger vehicles, having no idea that the people inside were studying hyenas. He joined the project in 2010 as an assistant cook, and his dedication and aptitude quickly became clear as he took on increasing responsibility within the field operation. By 2011 he had become a Research Assistant, and by 2020 he was leading field operations as Director of Operations.

Benson Pion as cook

2010

Dee White Joins the Project

Dee White and Kay Holekamp's connection goes back over 40 years. Dee was Kay's supervisor when Kay worked as a zookeeper at the St. Louis Zoo, long before the MHP existed. After Dee retired, she and Kay reconnected, and Dee began working as Kay's data manager and assistant. In 2010, Kay invited her to visit the field site in Kenya, fulfilling a lifelong dream of Dee's. She traveled to the Mara with Kay for many summers and later joined the project formally as Data Manager, where her meticulous care has helped maintain the integrity of one of the longest-running animal behavioral datasets in the world.

Dee White at the St. Louis Zoo
Dee White
Kay Holekamp at the St. Louis Zoo
Kay Holekamp

2009–2013

A New Generation Joins the MHP

Six young scientists who would go on to lead the MHP each began as volunteer Research Assistants in the Maasai Mara before pursuing their PhDs with Kay:

  • Kenna Lehmann, RA 2009–2010
  • Zach Laubach, RA 2010–2011
  • Tracy Montgomery, RA 2010–2011
  • Malit Benson Pion, RA 2011
  • Eli Strauss, RA 2011–2012
  • Lily Johnson-Ulrich, RA 2013–2014
Kenna Lehmann as RA Zach Laubach as RA Tracy Montgomery as RA Benson Pion as RA Eli Strauss as RA Lily Johnson-Ulrich as RA

2017–present

The Transition Team

Eli, Tracy, Kenna, Zach, and Lily form the "Transition Team" with the explicit goal of continuing the MHP past Kay's retirement, ensuring the long-term continuity of a study that cannot be replicated or restarted.

2018

30th Anniversary

The MHP marks 30 years of continuous operation, with over 150 peer-reviewed publications and a growing team of students and collaborators who trained with the project and have carried its questions forward into their own research programs.

Dr. Kay Holekamp

2020

Benson Becomes Director of Operations

Malit Benson Pion, who joined the project as an assistant cook in 2010 and worked as a Research Assistant from 2011, takes on the role of Director of Operations, leading the day-to-day running of both field camps and becoming an indispensable anchor of the entire project.

Benson Pion as Director of Operations

2023–2025

A Year of Transitions

Dr. Kenna Lehmann joins the faculty of Michigan State University in 2023. In 2025, Kay Holekamp retires from full-time faculty duties after 37 years at Michigan State University, though she continues to visit the field every summer and is currently writing a book about spotted hyenas. Dr. Eli Strauss and Dr. Tracy Montgomery join MSU as Assistant Professors, and Dr. Lily Johnson-Ulrich joins Hunter College, CUNY as an Assistant Professor.

Present

35+ Years and Counting

The MHP continues daily monitoring of seven hyena clans across the Maasai Mara, with an expanded team of Co-Directors, a dedicated Kenyan field staff, and new research initiatives on collective behavior, GPS collaring, cognition, urbanization, and conservation. The questions Kay and Laura started asking in 1988 are still very much alive.

A Legacy Built by Many

Hundreds of students, field assistants, and collaborators have contributed to the MHP over the decades. The project's continuity is a testament to the commitment of a remarkable team spread across two continents.

Meet the Team Our Mission
Hyena cubs in the Maasai Mara