Olivia Spagnuolo (Ph.D., 2024)
Eli Strauss (Ph.D., 2019)
Eli Swanson (Ph.D., 2013)
Micaela Szykman Gunther (Ph.D., 2001)
Jaime Tanner (Ph.D., 2007)
Kevin Theis (Ph.D., 2008)
Julie Turner (Ph.D., 2018)
Russ Van Horn (Ph.D., 2003)
Page Van Meter (Ph.D., 2009)
Aaron Wagner (Postdoctoral)
Sofia Wahaj (Ph.D., 2004)
Heather Watts (Ph.D., 2007)
​Kate Yoshida (Ph.D., 2012)

Joe Kolowski (Ph.D., 2007)
Suzanne LaCroix (Ph.D., 2011)
Zach Laubach (Ph.D., 2019)
Agathe Laurence ​(Postdoctoral)
Kenna Lehmann (Ph.D., 2020)
​Nora Lewin (Ph.D., 2017)
Kevin McCormick (Ph.D., 2022)
Terri McElhinny (M.S., 1996)
​Tracy Montgomery (Ph.D., 2020)
Eva Maria Muecke (Ph.D., 2005)
Scott Nunes (Ph.D., 1997)
Wiline Pangle (Ph.D., 2008)
​Connie Rojas (Ph.D., 2021)
Jenn Smith (Ph.D., 2010)

Sarah Benson-Amram (Ph.D., 2011)
Erin Boydston (Ph.D., 2001)
Katy Califf (Ph.D., 2013)
Leslie Curren (Ph.D., 2012)
Stephanie Dloniak (Ph.D., 2004)
Anne Engh (Ph.D., 2002)
Andy Flies (Ph.D., 2012)
Julia Greenberg (Ph.D., 2017)
David Green (Ph.D., 2015)
Keron Greene (M.S., 2024)
Timothy Ikime (M.Sc., 2015)
Julie Jarvey (Ph.D., 2024)
Lily Johnson-Ulrich (Ph.D., 2020)
Sarah Jones (Ph.D., 2019)

Since 1996, Dr. Kay Holekamp has mentored more than 35 students in pursuit of continued education. 

Previous Students

Dr. Kay Holekamp
Founder

history of the mara hyena Project

Founded in 1988, the Mara Hyena Project has been actively researching spotted hyenas in the Maasai Mara National Reserve. 

Meet Our Founder, Dr. Kay Holekamp

Kay founded the Mara Hyena Project with Dr. Laura Smale in 1988